Ancient 400-Mile Volcano Chain Buried Beneath China Reveals Secrets of Supercontinent Rodinia

A real scientific discovery of a ~700 km (about 400- mile) long fossilized Tonian- age magmatic (volcanic) arc belt buried beneath the Sichuan Basin in the Yangtze Block of South China, dating to roughly 820- 770 million years ago.

Rodinia was a Neoproterozoic supercontinent that existed roughly between 1.3- 0.75 billion years ago. It is the best- documented Precambrian supercontinent and played a central role in Earth’s tectonic, climatic, and biological evolution during the late Precambrian.

Illustration depicting the ancient 400-mile volcano chain buried beneath southern China, revealing sections of the supercontinent Rodinia.
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A real scientific discovery of a ~700 km (about 400- mile) long fossilized Tonian- age magmatic (volcanic) arc belt buried beneath the Sichuan Basin in the Yangtze Block of South China, dating to roughly 820- 770 million years ago.

Researchers from PetroChina and Nanjing University (and colleagues) combined:

  • Aeromagnetic surveys that revealed a NE- SW -trending positive magnetic anomaly belt (~700 km long and ~50 km wide), indicating iron- rich/(ultra)mafic rocks in the lower crust.
  • Seven deep boreholes (3.6- 6.5 km deep) that recovered Tonian (ca. 820- 770 Ma) rhyolites/granites of magmatic-arc affinity, overlain by Ediacaran sediments.
  • Geochronology, geochemistry, and seismic data confirming juvenile arc- related magmatism.

This places the belt well inland (at least 400- 900 km from the northwestern margin of the Yangtze Block), extending previously known circum-Yangtze Tonian arc systems (such as the Panxi- Hannan arc) into the continental interior. The preferred explanation is flat- slab subduction, in which an oceanic plate slides nearly horizontally far inland before steepening, producing magmatism far from the trench, analogous to processes observed in the modern Andes and inferred for other ancient systems.

The rocks formed during the Tonian period of the Neoproterozoic, around the time of the assembly and breakup of the supercontinent Rodinia. The structures have been dormant and buried under later sedimentary cover for hundreds of millions of years.

Broader implications

  • It expands understanding of Neoproterozoic tectonics and continental- arc systems in South China.
  • Large volcanic arcs can influence the global carbon cycle (via CO₂ release during eruptions and later drawdown through weathering), so this system is relevant to debates about Neoproterozoic climate, including possible contributions to “Snowball Earth” conditions, though the paper itself focuses primarily on the tectonic/magmatic evidence rather than claiming it rewrites climate history single-handedly.
  • It demonstrates how deep drilling and modern geophysics can reveal “hidden” ancient crustal architecture in stable basins that are otherwise inaccessible.

This Phys.org piece is a straightforward, accurate report of the peer- reviewed study (unlike some more sensational popular- science articles).

Geological Context of Rodinia

Rodinia was a Neoproterozoic supercontinent that existed roughly between 1.3- 0.75 billion years ago. It is the best- documented Precambrian supercontinent and played a central role in Earth’s tectonic, climatic, and biological evolution during the late Precambrian.

Timeline

PhaseApproximate AgeKey Events
Assembly1300- 900 MaGlobal mountain- building (Grenville- age orogenies). Continents accreted mainly around Laurentia (proto- North America).
Stable existence~900- 825 MaSupercontinent largely intact for ~150 million years.
Early rifting / breakup initiation825- 750 MaMantle superplume activity, widespread rifting, and bimodal magmatism. Major pulses at ~825 Ma, ~780 Ma, and ~750 Ma.
Main breakup750- 600 MaContinents dispersed diachronously. Opening of the proto-Pacific and other oceans.
Aftermath~630- 530 MaFragments reassembled into the short- lived Pannotia / greater Gondwana.

Configuration

Most reconstructions place Laurentia at the center of Rodinia. Other major blocks (Baltica, Amazonia, Australia, East Antarctica, India, Siberia, Congo, Kalahari, South China, etc.) were arranged around it. Exact positions remain debated, especially for South China:

  • Older “Missing Link” models placed South China between Laurentia and Australia.
  • Many recent models favor a peripheral (external) position, often near Australia- India- East Antarctica.

Tectonic & Geodynamic Significance

  • Rodinia’s assembly involved long- lived accretionary orogens that eventually collided (Grenville, Sveconorwegian, etc.).
  • Its breakup is widely linked to a mantle superplume (or superswell) that developed beneath the supercontinent due to thermal insulation and slab avalanches. This triggered large igneous provinces, rifting, and eventually continental separation.
  • The breakup created vast new passive margins and ocean basins, dramatically altering global geography.

Climatic & Biological Context

The breakup of Rodinia coincided with (and likely helped trigger) extreme climate events:

  • The Cryogenian “Snowball Earth” glaciations (Sturtian ~720- 660 Ma and Marinoan ~650- 635 Ma).
  • Dramatic changes in the carbon cycle, ocean chemistry, and the eventual rise of complex multicellular life in the Ediacaran and Cambrian.

Link to the Sichuan Basin Discovery

The ~700 km fossilized magmatic arc (820- 770 Ma) under China’s Sichuan Basin formed during the early breakup phase of Rodinia. It records subduction (likely flat- slab) along or near the margin of the Yangtze Block while the supercontinent was beginning to rift apart. This makes the discovery a direct window into the tectonic processes operating at the edges of Rodinia ~800 million years ago.

In short, Rodinia represents a pivotal chapter in Earth’s history, the last major supercontinent before the modern plate- tectonic regime fully matured and complex life exploded onto the scene.

A ∼700-km-Long Fossil Tonian Magmatic Arc Belt Hidden Within the Yangtze Block’s Interior, South China

Scientists have identified a ~700 km (about 400- mile) long fossilized magmatic arc (a chain of ancient volcanoes) buried deep beneath the Sichuan Basin in the interior of the Yangtze Block, South China.

  • It dates to the Tonian period of the Neoproterozoic era, specifically ca. 820- 770 million years ago.
  • The arc is marked by an iron- rich strip of rock detected through aeromagnetic surveys.
  • Confirmation came from seven deep boreholes (3.6- 6.5 km deep) that recovered rhyolites and granites of clear magmatic -arc chemical affinity.
  • The belt stretches inland at least 400- 900 km from the northwestern edge of the Yangtze Block, much farther inland than typical volcanic arcs.

The researchers attribute the unusually inland position to flat- slab subduction. In this process, an oceanic plate slides nearly horizontally beneath the continent for a long distance before sinking more steeply into the mantle. This can create magmatism far from the continental margin.

The new arc is interpreted as an inland extension of the known Panxi- Hannan arc system and is linked to southeastward oceanic subduction during the assembly and breakup of the supercontinent Rodinia.

Abstract and Core Findings

Fossil magmatic arcs are important records of juvenile magmatism, crustal growth, and differentiation. Some ancient arcs, however, lie buried in continental interiors under thick sedimentary cover, making them hard to detect.

This study uses a combination of:

  • Seven deep boreholes (∼3,600- 6,500 m deep) that reached the basement of the Sichuan Basin
  • Geochronology
  • Geochemistry
  • Aeromagnetic data
  • 3D seismic reflection profiles

Key results:

  • Widespread Tonian (ca. 820- 770 Ma) rhyolites and granites occur beneath the central Sichuan Basin, directly overlain by Ediacaran sediments.
  • Aeromagnetic surveys reveal a NE- SW- trending ∼700- km- long positive magnetic anomaly belt (PMAB) across the central basin. This indicates the presence of (ultra-) mafic rocks in the lower crust.
  • The felsic rocks recovered from boreholes within the PMAB show geochemical signatures consistent with derivation from a juvenile arc basaltic source.

The authors interpret the PMAB as recording the input of mantle- derived melts and their subsequent differentiation into granitic magmas. This represents a Tonian arc magmatic front that once existed in the interior of the Yangtze Block.

Tectonic Implications

  • The discovery extends the known circum-Yangtze Tonian continental arc system (including the Panxi-Hannan arc) far into the Yangtze Block’s interior.
  • The arc reaches at least 400- 900 km landward from the northwestern margin of the Yangtze Block.
  • The preferred explanation is flat- slab subduction, which allows magmatism to occur unusually far inland.

Significance

This is one of the best- documented examples of a long, inland fossil magmatic arc of Tonian age. It improves our understanding of the tectonic- magmatic evolution of South China during the breakup of the supercontinent Rodinia and shows how deep drilling and modern geophysics can reveal previously hidden crustal architecture.

Title: A ∼700‐km‐Long Fossil Tonian Magmatic Arc Belt Hidden Within the Yangtze Block’s Interior, South China

Published:  Journal of Geophysical Research: Solid Earth (2025)

DOI: 10.1029/2024JB030825

Authors: Zhidong Gu, Junyong Li, Xiaolei Wang, Ya Xu, Xiufen Zhai et al.

Abstract

Fossil magmatic arcs preserve important archives of juvenile magmatism, crustal growth and differentiation, thereby aiding in our understanding of continental evolutionary histories. However, some ancient arcs are situated in continental interiors, where they are buried beneath thick cover sequences, making their identification challenging. Here, we present a data set of deep boreholes, geochronology, geochemistry and geophysics to investigate the basement properties and evolution of the Sichuan Basin in the Yangtze Block, South China. These results provide evidence for a large Tonian magmatic arc belt hidden within the Yangtze Block’s interior. Seven deep boreholes (∼3,600–6,500 m) penetrating into basin basement, together with geochronology and 3D seismic reflection profiles, show extensive Tonian (ca. 820–770 Ma) rhyolite/granites overlain by Ediacaran sediments. Aeromagnetic data delineate a NE-SW-trending, ∼700-km-long positive magnetic anomaly belt (PMAB) across the central basin, indicating the presence of (ultra-) mafic rocks in lower crust. Petrogenetic analyses indicate that the Tonian felsic rocks drilled within PMAB range were likely derived from a juvenile arc basaltic source. Thus, we interpret the PMAB recorded input of mantle-derived melts and their subsequent evolution into granitic magmas, associated with a Tonian arc magmatic front that once occurred in the Yangtze Block’s interior. Our finding broadens the range of the Tonian continental arc system in the region, reaching at least 400–900 km landward from block’s northwestern margin. Such a broad orogenic system played a vital role in continental evolution, element recycling and basin formation of the Yangtze Block.

Gondwana Was a True Supercontinent—and It Helped Spark the Explosion of Life on Earth

Gondwana (also called Gondwanaland) was a vast southern supercontinent that existed from the late Neoproterozoic through much of the Paleozoic and into the Mesozoic. It is one of the best- documented ancient landmasses in Earth’s history.

The Cambrian explosion (roughly 541- 520 million years ago, peaking around 538- 520 Ma) marks the relatively rapid appearance and diversification of most major animal body plans (phyla) in the fossil record, including arthropods, molluscs, chordates, echinoderms, and others.

An artistic representation of Gondwana, the supercontinent, 550 million years ago, featuring mountainous landscapes, volcanic activity, and marine life from the Cambrian period.
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Gondwana (also called Gondwanaland) was a vast southern supercontinent that existed from the late Neoproterozoic through much of the Paleozoic and into the Mesozoic. It is one of the best- documented ancient landmasses in Earth’s history.

The name comes from the Gondwana region of central India.

Austrian geologist Eduard Suess first proposed the concept in 1885. He noted the striking similarity of fossil plants (especially the Glossopteris flora) and geological sequences across India, Australia, Africa, Antarctica, and South America, concluding these lands were once joined.

Alfred Wegener later incorporated Gondwana into his 1912 continental- drift hypothesis as the southern counterpart to Laurasia within the larger supercontinent Pangaea.

Gondwana assembled over a prolonged period through the collision of several older continental blocks (cratons) during the late Neoproterozoic to early Cambrian:

  • Major collisions occurred roughly between 650 and 500 million years ago.
  • Key events included the East African Orogeny and related mountain- building episodes that sutured West Gondwana (Africa and South America) to East Gondwana (India, Antarctica, Australia and Madagascar).
  • By about 550- 500 Ma, the core of Gondwana was largely complete.

Recent research (2026) significantly expands this picture. Using samarium- neodymium isotope mapping of thousands of granite samples, scientists identified previously unrecognized crustal fragments (now buried under younger mountains, especially across large parts of Asia from India through China into Kazakhstan) that belonged to Gondwana.

This “Greater Gondwana” is estimated to have comprised roughly 80% of Earth’s continental landmass around 550- 500 Ma, qualifying it as a true supercontinent under common definitions. Its assembly and the subsequent formation of a vast volcanic arc (a “Cambrian ring of fire”) are now linked to major climate shifts and the Cambrian explosion of complex life.

Life of the Supercontinent

  • During the Paleozoic, Gondwana drifted as a coherent unit, often positioned near or over the South Pole at various times.
  • It experienced extensive glaciation in the late Paleozoic (the great Carboniferous- Permian ice age).
  • In the late Paleozoic it collided with Laurasia (North America, Europe and Asia) to form the even larger supercontinent Pangaea (~300- 250 Ma). Gondwana formed the southern half of Pangaea.

Breakup

Gondwana began to fragment in the Mesozoic as Pangaea itself broke apart:

  • Initial rifting started in the Early Jurassic (~180 Ma).
  • The sequence of separation roughly followed this order:
    • India began separating and drifted northward.
    • Africa and South America split (opening the South Atlantic).
    • Australia and Antarctica separated later.
    • Madagascar and other smaller fragments followed.
  • By the Late Cretaceous to early Cenozoic, the modern southern continents and the Indian subcontinent were largely independent.

The former pieces of Gondwana are today the continents of South America, Africa, Antarctica, Australia, the Indian subcontinent, Madagascar, and numerous smaller terranes and microcontinents.

Geological, paleontological, and paleomagnetic evidence continues to refine the exact configuration and timing of its assembly and dispersal.

The 2026 “Greater Gondwana” findings have renewed interest in how its size and tectonic activity influenced global climate, ocean chemistry, and the rise of complex animal life during the Cambrian.

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Gondwana really was a supercontinent—and it may have helped kickstart life on Earth, by Bill Collins (Adjunct Professor, School of Earth and Planetary Sciences, Curtin University), published in The Conversation on 9 September 2026.

The article discusses a new Science Advances paper coauthored by Collins. It argues that Gondwana (formed roughly 550- 500 million years ago) qualified as a true supercontinent and that its formation helped drive the Cambrian explosion of complex life.

Around 550 million years ago, the Southern Hemisphere was dominated by Gondwana, first recognized in 1885 by Austrian geologist Eduard Suess from matching plant fossils across India, Australia, Africa, Antarctica, and South America.

Supercontinents are defined in the context of cyclic, global- scale sea- level changes going back at least 2 billion years. Recognized ones include Pangaea (youngest, ~300- 250 Ma), Gondwana, Rodinia (~1 Ga), and Columbia/Nuna (~2- 1.6 Ga).

Gondwana’s supercontinent status was debated because estimates put it at only ~64% of continental landmass (below an arbitrary 75% threshold often used). Even adding fragments from the Appalachians through southern Europe, Turkey, Iran, and Pakistan left it short.

The team serendipitously discovered buried Gondwanan crust while compiling a global granite dataset to study deep crustal processes and mineral formation. They used samarium- neodymium isotope analysis to map the age of source rocks for surface granites at a continental scale.

  • Example: Himalayan granites of various ages (20 Ma, 50 Ma, 120 Ma) all derive from a common continental block that formed ~700- 550 Ma and belonged to Gondwana.
  • They traced original Gondwanan fragments from India and Southeast Asia through most of China into Kazakhstan. Much of this region (now mountainous and covered by younger rocks) was previously thought to be ancient ocean floor with volcanic island chains.
  • This expands Gondwana to roughly 80% of Earth’s continental landmass at the time, confirming supercontinent status.

Supercontinent formation affects global climate and biology.

After greater Gondwana assembled (between ~750- 550 Ma), Himalayan- scale mountain ranges crossed the frozen, arid landmass. Plate tectonics then reconfigured, forming a massive volcanic arc (a “Cambrian ring of fire”) nearly 35,000 km long that encircled the supercontinent, from Russia via China, Australia, Antarctica, and South Africa to the Andes. This was a precursor to today’s Pacific Ring of Fire.

Constant eruptions released large volumes of volcanic gases (especially water vapor and CO₂).

Over ~50 million years, these emissions shifted Earth’s climate from icehouse to greenhouse conditions. The authors argue this global change was a key catalyst for the Cambrian explosion (~550- 500 Ma), when complex life proliferated.

Other coverage of the same study (e.g., in Science, Scientific American, and 404 Media) notes the analysis of >25,000 rock samples and links the expanded Gondwana to effects on sea level, atmospheric gases, volcanism, and early life.

Some external experts accept a larger Gondwana but note potential implications for reconstructions of later supercontinents like Pangaea.

The full article is available at: https://theconversation.com/gondwana-really-was-a-supercontinent-and-it-helped-kick-start-life-on-earth-291460.

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The Cambrian explosion (roughly 541- 520 million years ago, peaking around 538- 520 Ma) marks the relatively rapid appearance and diversification of most major animal body plans (phyla) in the fossil record, including arthropods, molluscs, chordates, echinoderms, and others.

Complex multicellular life with skeletons, mobility, predation, and sophisticated ecosystems emerged after billions of years dominated by simpler forms (microbes and Ediacaran biota).

It was not a single instantaneous event but a cascade of radiations spanning the late Ediacaran into the early Cambrian, with pulsed increases in diversity, disparity (range of body forms), biomineralization, and ecological complexity.

No single cause fully explains it; instead, interacting intrinsic (genetic/developmental), extrinsic (environmental/geological), and ecological factors created permissive conditions and feedback loops.

1. Environmental and Geological Triggers

These set the timing and enabled higher metabolic rates, larger sizes, and more active lifestyles.

Rising and fluctuating oxygen levels: Oxygen had been increasing stepwise since the Great Oxidation Event but further rises (or expansions of oxic habitats) in the late Neoproterozoic- early Cambrian crossed thresholds allowing larger, more energetic animals and carnivory. Even modest levels (a few percent of present atmospheric levels) may have sufficed for early animals; modern analogs in oxygen minimum zones show carnivores are rare under very low oxygen. Dynamic redox conditions and oxygenation pulses coincided with biodiversity increases.

Nutrient availability and ocean chemistry: Enhanced continental weathering (from mountain building and volcanism) delivered phosphorus, calcium, and other elements. This boosted primary productivity (algae/phytoplankton), supported food webs, and enabled biomineralization (shells, exoskeletons, skeletons using Ca and other minerals). Sea- level rises flooded continents, creating extensive shallow marine habitats and new niches.

Tectonics, volcanism, and climate shifts: Assembly of the supercontinent Gondwana (~550- 500 Ma) involved major mountain building, followed by reconfiguration of plate tectonics. This created a vast volcanic arc (“Cambrian ring of fire,” ~35,000 km long, precursor to today’s Pacific Ring of Fire) that released large amounts of CO₂ and water vapor over tens of millions of years. This helped shift climate from icehouse (post- Snowball Earth) conditions toward a warmer greenhouse state, altering ocean chemistry and promoting life’s expansion. Recent work linking expanded Gondwana (~80% of continental landmass) strengthens this connection between deep- Earth processes and the explosion. Earlier studies also tied Gondwana-related volcanism to temperature rises and evolutionary bursts. Sea- level changes from rifting and mid- ocean ridge activity further contributed by expanding habitats.

Other factors include carbon- cycle perturbations (isotope excursions reflecting productivity and anoxia changes) and possible orbital forcing of nutrient pulses.

2. Ecological Mechanisms

Once environmental thresholds were crossed, biotic interactions accelerated diversification:

Predator- prey arms races: The rise of carnivory (enabled by oxygen) created selective pressure for defenses (hard skeletons, spines, mobility, burrowing) and sensory systems (e.g., eyes). This escalated morphological innovation and complexity. Evidence includes early predatory traces, drill holes, and defensive morphologies.

Ecosystem engineering and feedbacks: Animals began bioturbating sediments (mixing them), clearing microbial mats, and altering nutrient cycles and redox conditions. This created more complex food webs, new niches, and positive feedback that sustained further radiation. Suspension feeding may have clarified water columns, aiding photosynthesis.

The result was a shift from relatively simple Ediacaran communities to more structured, niche- partitioned Cambrian ecosystems with higher trophic levels.

3. Developmental and Genetic Foundations

These provided the capacity for novelty but were not the sole trigger (many key gene families predate the Cambrian):

  • Evolution of gene regulatory networks (including Hox genes) allowed modular body plans, segmentation, bilateral symmetry, and appendages.
  • Developmental flexibility enabled rapid experimentation with forms once ecological opportunities opened.
  • Molecular clocks indicate some animal lineages originated earlier (deep “fuse”), with the fossil explosion reflecting when they became abundant, mineralized, and preserved.

Most researchers favor a multifactorial model: geological/tectonic changes (including Gondwana assembly and associated volcanism) drove environmental shifts (oxygen, nutrients, climate, habitats). These crossed critical thresholds enabling active lifestyles and predation. Ecological interactions then produced rapid morphological and taxonomic diversification via arms races and niche filling, all built on preexisting genetic toolkits.

The event is unique in the scale of body- plan innovation; later radiations (e.g., Ordovician) increased diversity within existing plans.

Ongoing research continues to refine timings, redox records, and tectonic links, but the consensus is that the Cambrian explosion resulted from the coincidence of permissive environments and evolutionary opportunities rather than any isolated cause.

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Buried fragments reveal a “Greater Gondwana” supercontinent

The study reconstructs Greater Gondwana as a true supercontinent that encompassed roughly 80% of Earth’s continental landmass around 550- 500 million years ago (centered near the South Pole). This exceeds the commonly cited (though arbitrary) ~75% threshold for supercontinent status. Earlier estimates placed classic Gondwana (Africa, Antarctica, Australia, India, South America, plus some extensions) at about 64%.

Previously unrecognized (or reclassified) crustal fragments now buried under younger mountains, particularly in Asia (from India/Southeast Asia through much of China into Kazakhstan), with implications for parts of Europe and elsewhere, belonged to this expanded landmass. Many of these areas were previously interpreted as ancient ocean floor or volcanic island arcs.

Researchers compiled a global dataset of >25,000 granite/igneous rock samples. They used samarium-neodymium (Sm-Nd) isotope systematics to determine the model ages of the deep crustal sources from which the granites were derived (most granites form by melting older continental crust).

Mapping these ages at continental scale revealed coherent blocks that formed ~700- 550 Ma and shared a common history with Gondwana, even where surface rocks are much younger (e.g., Himalayan granites of various ages all tracing back to the same ~700- 550 Ma block). This approach “sees through” later mountain- building events.

The discovery was described as serendipitous; the team was originally studying deep crustal processes and mineralization.

Gondwana’s assembly involved major orogeny (mountain building). After amalgamation, plate tectonics reconfigured, producing a nearly continuous volcanic arc (“Cambrian ring of fire,” ~35,000 km long) encircling the supercontinent, a precursor to the modern Pacific Ring of Fire.

Prolonged volcanism released large volumes of greenhouse gases (CO₂ and water vapor), helping shift global climate from icehouse (post- Snowball Earth) to greenhouse conditions over ~50 million years. Combined with effects on sea level, nutrient delivery (via enhanced weathering), atmospheric composition, and ocean chemistry, this is proposed as a major catalyst for the Cambrian explosion of complex animal life (~538- 500 Ma). The paper links deep- Earth tectonic processes to this profound biological radiation.

Context and Reception

  • Supercontinent definitions remain somewhat debated; the 75% figure is not absolute.
  • External commentary notes that assigning these Asian blocks to Gondwana may require rethinking later configurations (e.g., Pangaea), though the fragments may have later drifted away.
  • The work strengthens connections between Neoproterozoic- Cambrian tectonics, climate, and the rise of complex life.

Full text access may require institutional subscription or purchase via the Science Advances site. Summaries and commentary appear in The Conversation (by co- author Collins), Science, Scientific American, 404 Media, and other outlets.

Published: Science Advances

DOI: 10.1126/sciadv.aeh6155

Provided: The Conversation

Authors: Tao Wang (Chinese Academy of Geological Sciences),

William J. Collins,

Chaoyang Wang,

Zengqian Hou,

Ying Tong,

He Huang

and Jieping Ye

Abstract

Profound global biogeochemical shifts marked the Ediacaran-Cambrian transition [550–500 million years (Ma)], a phenomenon often ascribed to the dynamics of supercontinent cycles. However, Gondwana is frequently denied supercontinent status due to its perceived spatial deficit (∼64% of continental landmass). Leveraging continental-scale Nd isotopic mapping based on a vast database of 25,346 felsic-intermediate igneous rocks, we reveal extensive buried Precambrian terranes beneath Phanerozoic orogens in Asia, Europe, and North America. Our results expand Gondwana’s reconstructed area to ∼80% of the global landmass, firmly establishing its identity as a true supercontinent—termed “Greater Gondwana.” We propose that the assembly and peripheral subduction of this “Greater Gondwana” played a pivotal role in driving the geodynamic and environmental perturbations that culminated in the Cambrian metazoan radiation.